Blue roses were traitionally created by dyeing white roses, since roses lack a gene to produce delphinidin, the primary plant pigment that produces true blue flowers. So-called "blue roses" have been bred by conventional hybridization methods, but the results, such as "Blue Moon" are more accurately described as lilac in color. However, after thirteen years of joint research by an Australian company Florigene, and Japanese company Suntory, a blue rose was created in 2004 using genetic engineering. The delphinidin gene was cloned from the petunia and inserted into a mauve-blend rose, the Old Garden Rose 'Cardinal de Richelieu' (a Rosa gallica). However, since the pigment cyanidin was still present, the rose was more dark burgundy than true blue. Further work on the rose using RNAi technology to depress the production of cyanidin produced a mauve colored flower, with only trace amounts of cyanidin.