The Five-Spotte Hawkmoth (Manduca quinquemaculata) is a brown and gray hawk moth of the Sphingidae family. The caterpillar is often referred to as the tomato hornworm and can be a major pest in gardens. Tomato hornworms are closely related to (and sometimes confused with) the tobacco hornworm (Manduca sexta). This confusion arises because caterpillars of both species feed on the foliage of various plants from the family Solanaceae, so either species can be found on tobacco or tomato leaves, and the plant on which the caterpillar is found does not indicate its species. The larva of these species can be distinguished by their lateral markings; tomato hornworms have eight V-shaped markings while tobacco hornworms have seven diagonal lines . Furthermore, the caterpillars can be distinguished from the larval stage onwards by the color of the horns on their back ends: M. quinquemaculata caterpillars have black horns, while Manduca sexta caterpillars have red horns. In addition, a small percentage of M. quinquemaculata caterpillars are black, whereas a black M. sexta caterpillar has not been observed in nature. The moths can be distinguished by the number of spots on their abdomen, with M. quinquemaculata having five.